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1995-1999
MSci in Chemistry at University College London

1999-2000
MRes in Molecular Functions in Disease at the University of Glasgow

2000-2003
PhD in the Monckton laboratory at the University of Glasgow – “Modelling the Toxic RNA Hypothesis of Myotonic Dystrophy in Drosophila Melanogaster”

2004-2006
Post-doc in the Tollervey laboratory at the Wellcome Centre for Cell Biology in Edinburgh, studying RNA degradation in budding yeast.

2006-2008
Leverhulme Trust Early Career Fellowship in the Tollervey laboratory at the Wellcome Centre for Cell Biology in Edinburgh, “Substrate recognition by the TRAMP complexes”

2008-2009
Post-doc in the Tollervey laboratory at the Wellcome Centre for Cell Biology in Edinburgh, studying rDNA recombination and non-coding RNA biology.

2009-2014
Wellcome Trust Research Career Development Fellow at the ����������̳

2014-2016
Group leader at the ����������̳

2016-present
Wellcome Trust Senior Research Fellow at the ����������̳